| Main Ref. | Holthuis, L.B., 1980 | 
|---|---|
| Remarks | Juveniles are found in mangrove creek-inlets (Ref. 121464), in a mangrove vegetate area (Ref. 121475), in intertidal and subtidal edge of mud flats, near inshore and far inshore waters (Ref. 121464). Occurs in bottom mud or sandy-mud substrates in marine and estuarine environments (Ref. 8) and among shells (Ref. 121467). Inhabits shallow open sea or in the mouth of a river and bay areas where water is more or less turbid (Ref. 374). Adult species periodically form aggregations or 'schools' offshore (Ref. 100847). Omnivore (Ref. 116259). | 
| Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
|---|---|---|---|---|
| Marine zones / Brackish and freshwater bodies | 
 | 
 | 
 | 
 | 
| Substrate | Benthic: mobile; Soft Bottom: sand; mud; | 
|---|---|
| Substrate Ref. | del Mundo, C.M., 2000 | 
| Special habitats | Beds: sea grass; | 
| Special habitats Ref. | Hajisamae, S. and P. Yeesin, 2014 | 
| Ref. | Holthuis, L.B., 1980 | 
|---|---|
| Associations | |
| Associated with | |
| Association remarks | |
| Parasitism | 
| Feeding type | mainly animals (troph. 2.8 and up) | 
|---|---|
| Feeding type Ref. | Wassenberg, T.J. and B.J. Hill, 1993 | 
| Feeding habit | hunting macrofauna (predator) | 
| Feeding habit Ref. | Wassenberg, T.J. and B.J. Hill, 1993 | 
| Estimation method | original sample | unfished population | Remark | ||
|---|---|---|---|---|---|
| Troph | s.e. | Troph | s.e. | ||
| From diet composition | |||||
| From individual food items | 3.77 | 0.36 | Trophic level estimated from a number of food items using a randomized resampling routine. | ||
| Ref. | |||||