Jefferson, T.A., S. Leatherwood and M.A. Webber. 1993. (Ref. 1394)
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Climate / Range
Subtropical; 45°N - 13°N, 140°W - 180°E
Dusky black with lighter ventrum. Gray head and shoulders, black nose with brown-red hair around the eyes. Guard hairs are white-tipped in adult males. Claws present in 5 fingers and absent on the 1st and 5th toes. Cartilaginous webbing extends past the claws. Females have 2 pairs abdominal nipples.
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Estimation of some characteristics with mathematical models