| Main Ref. | Holthuis, L.B., 1991 |
| Mode | |
| Fertilization | |
| Spawning Frequency | one clear seasonal peak per year |
| Batch Spawner | No |
| Reproductive Guild | |
| Description of life cycle and mating behavior | Breeds once a year between June and October. Larger females mate and lay eggs earlier during breeding season while smaller females spawn towards the end (Ref. 85364). |
| Search for more references on reproduction | Scirus |