| Citation | Bisby, F., Y. Roskov, A. Culham, T. Orell, D. Nicolson, L. Paglinawan, N. Bailly, W. Appeltans, P. Kirk, T. Bourgoin, G. Baillargeon and D. Ouvrard 2012 Species 2000 & IT IS Catalogue of Life: 2012 Annual Checklist. Digital resource at www.catalogueoflife.org/annual-checklist/2012/. Species 2000: Reading, UK. |
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| Ref. No. | 91944 |
| Language | English |
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